A. Phylogeny-group-;-?-ashelford, K. E. Chuzhanova, N. A. Fry, J. C. Jones, A. J. Weightman et al., At least 1 in 20 16S rRNA sequence records currently held in public repositories is estimated to contain substantial anomalies, Botanical Journal of the Linnean Society, vol.181, pp.7724-7736, 2005.

J. F. Basinger and D. C. Christophel, Fossil flowers and leaves of the Ebenaceae from the Eocene of southern Australia, Canadian Journal of Botany, vol.63, pp.1825-1843, 1985.

J. F. Basinger, D. R. Greenwood, P. G. Wilson, and D. C. Chistophel, Fossil flowers and fruits of capsular Myrtaceae from the Eocene of South Australia, Canadian Journal of Botany, vol.85, pp.204-215, 2007.

?. Batten, D. Bell, C. D. Soltis, D. E. Soltis, P. Boucher et al., An extinct genus of Salicaceae based on twigs with attached flowers, fruits, and foliage from the Eocene Green River Formation of Utah and Colorado, Stratigraphy, palaeogeography and evolutionary significance of Late Cretaceous and Early Tertiary Normapolles pollen, vol.35, pp.1389-1399, 1981.

R. Bouckaert, T. G. Vaughan, J. Barido-sottani, S. Duchêne, M. Fourment et al., BEAST 2.5: An advanced software platform for Bayesian evolutionary analysis, PLoS Computational Biology, vol.15, p.1006650, 2019.

B. Boyle, N. Hopkins, Z. Lu, R. Garay, J. Mozzherin et al., Investigations of angiosperms from the Eocene of southeastern North America: Samaras of Fraxinus wilcoxiana Berry, Review of Palaeobotany and Palynology, vol.14, pp.1078-1083, 1992.

S. Chamberlain, E. Szocs, C. Boettiger, K. Ram, I. Bartomeus et al., Seed morphology of modern and fossil Ampelocissus (Vitaceae) and implications for phytogeography, The importance of phylogenetic structure in biodiversity studies, vol.95, pp.1534-1553, 2007.

M. J. Christenhusz and J. W. Byng, The number of known plants species in the world and its annual increase, Phytotaxa, vol.261, pp.201-217, 2016.

R. A. Christopher, Normapolles and triporate pollen assemblages from the Raritan and Magothy formations (upper Cretaceous) of New Jersey, Palynology, vol.3, pp.73-122, 1979.

T. L. Couvreur, M. D. Pirie, L. W. Chatrou, R. M. Saunders, Y. C. Su et al., A preliminary survey of fossil leaves and well-preserved reproductive structures from the Sentinel Butte Formation (Paleocene) near Almont, North Dakota, Journal of Biogeography, vol.38, pp.328-344, 1990.

P. R. Crane, E. M. Friis, K. Pedersen, P. R. Crane, E. M. Friis et al., Paleobotanical evidence on the early radiation of magnoliid angiosperms, The anther: form, function and phylogeny, vol.8, pp.25-27, 1994.

W. L. Crepet and K. C. Nixon, Fossil Clusiaceae from the Late Cretaceous (Turonian) of New Jersey and implications regarding the history of bee pollination, American Journal of Botany, vol.85, pp.1122-1133, 1998.

?. Cronquist, A. Dagallier, L. Janssens, S. Dauby, G. Blach-overgaard et al., Cradles and museums of generic plant diversity across tropical Africa, Journal of Biogeography, 1981.

C. P. ?-daghlian, A review of the fossil record of monocotyledons, Botanical Review, vol.47, pp.517-555, 1981.

D. Dilcher and P. Crane, Archaeanthus: an early angiosperm from the Cenomanian of the Western Interior of North America, Annals of the Missouri Botanical Garden, vol.71, pp.351-383, 1984.

J. A. Doyle, C. L. Hotton, J. Ward, J. A. Doyle, S. Manchester et al., Early Cretaceous tetrads, zonasulculate pollen, and Winteraceae. 1. Taxonomy, morphology, and ultrastructure, American Journal of Botany, vol.77, pp.636-646, 1990.

?. Edelman, D. Moscow, U. Id, E. Estrada-ruiz, L. Calvillo-canadell et al., The Eocene Germer Basin flora of south-central Idaho, Taxonomic affinities of Medusagyne oppositifolia (Medusagynaceae), vol.90, pp.111-120, 1975.

J. ?-felsenstein, Phylogenies and quantitative characters, Annual Review of Ecology and Systematics, vol.19, pp.445-471, 1988.

X. Feng, B. Tang, T. M. Kodrul, and J. J. , Winged fruits and associated leaves of Shorea (Dipterocarpaceae) from the Late Eocene of South China and their phytogeographic and paleoclimatic implications, American Journal of Botany, vol.100, pp.574-581, 2013.

E. Friis, K. R. Pedersen, P. Crane, E. Friis, K. Pedersen et al., Endressianthus, a new Normapollesproducing plant genus of Fagalean affinity from the Late Cretaceous of Portugal, Fossil evidence of water lilies (Nymphaeales) in the Early Cretaceous, vol.410, pp.48-52, 1977.

C. Graham, P. Fine, P. ;-?-herendeen, W. Crepet, and D. Dilcher, Testing the spatial phylogenetic structure of local communities: statistical performances of different null models and test statistics on a locally neutral community, The fossil history of the Leguminosae: phylogenetic and biogeographic implications, vol.11, pp.303-316, 1992.

E. ?-hermsen, M. A. Gandolfo, K. C. Nixon, and W. L. Crepet, Divisestylus gen. nov. (aff. Iteaceae), a fossil saxifrage from the Late Cretaceous of, American Journal of Botany, vol.90, pp.1373-1388, 2003.

L. J. Hickey, A. Peterson, K. W. Hilu, T. Borsch, K. Müller et al., Synthesis of phylogeny and taxonomy into a comprehensive tree of life, Evolutionary dynamics and biogeography of Musaceae reveal a correlation between the diversification of the banana family and the geological and climatic history of Southeast Asia, vol.56, pp.1453-65, 1978.

S. Jardiné and H. Magloire, Palynologie et stratigraphie du Crétacé des Bassin du Sénégal et Cote d'Ivoire. Mémoires du Bureau de Recherches Géologiques et Miniéres, vol.32, pp.187-245, 1965.

D. M. ?-jarzen, Some Maastrichtian palynomorphs and their phytogeographical and paleoecological implications, Palynology, vol.2, pp.29-38, 1978.

K. Katoh, K. Misawa, K. Kuma, and T. Miyata, MAFFT: a novel method for rapid multiple sequence alignment based on Fourier transform, Nucleic Acids Research, vol.30, pp.3059-3066, 2002.

?. Kedves, M. Kissling, W. Knobloch, E. Mai, D. Kress et al., Has frugivory influenced the macroecology and diversification of a tropical keystone plant family? Research Ideas and Outcomes 3: e14944, Systematics, and Fossil History of the Hamamelidae, 1-7. Systematics Association Special Volume, 40b, vol.47, pp.8369-8374, 1986.

E. Lee, A. Cibrian-jaramillo, S. Kolokotronis, M. Katari, A. Stamatakis et al., A Functional Phylogenomic View of the Seed Plants, 2011.

D. P. ?-little and D. S. Barrington, Major evolutionary events in the origin and diversification of the fern genus Polystichum (Dryopteridaceae), American Journal of Botany, vol.90, pp.508-514, 2003.

Y. Li, T. Smith, C. J. Liu, N. Awasthi, J. Yang et al., Endocarps of Prunus (Rosaceae: Prunoideae) from the early Eocene of Wutu, vol.60, pp.555-564, 2011.

R. Lupia, S. Lidgard, and P. R. Crane, Comparing palynological abundance and diversity: Implications for biotic replacement during the Cretaceous angiosperm radiation, Paleobiology, vol.25, pp.305-340, 1999.

S. ?-magallón and A. Castillo, Angiosperm diversification through time, American Journal of Botany, vol.96, pp.349-365, 2009.

S. ?-magallón, A review of the effect of relaxed clock method, long branches, genes, and calibrations in the estimation of angiosperm age, Botanical Sciences, vol.92, pp.1-22, 2014.

S. ?-magallón, S. Gomez-acevedo, and L. L. Sanches-reyes, A metacalibrated time-tree documents the early rise of flowering plant phylogenetic diversity, New Phytologist, vol.207, pp.437-453, 2015.

D. H. Mai, Entwicklung der Wasser-und Sumpfpflanzen-Gesellschaften Europas von der Kreide bis ins Quartär, Feddes Repertorium, vol.176, issue.17, pp.197-229, 1985.

S. Manchester, J. Kress, S. Manchester, D. Kapgate, and J. Wen, Fossil bananas (Musaceae): Ensete oregonense sp. nov. from the Eocene of western North America and its phytogeographic significance, Annals of the Missouri Botanical Garden, vol.80, pp.1849-1859, 1993.

?. Martinez-millan, M. Crepet, W. L. Nixon, and K. C. , Pentapetalum trifasciculandricus gen. et sp. nov., a thealean fossil flower from the Raritan Formation, American Journal of Botany, vol.96, pp.933-949, 2009.

B. A. Mohr and M. E. Bernardes-de-oliveira, Endressinia brasiliana, a magnolialean angiosperm from the Lower Cretaceous Crato Formation (Brazil), International Journal of Plant Sciences, vol.165, pp.1121-1133, 2004.

M. Moore, C. Bell, P. Soltis, and D. Soltis, Using plastid genome-scale data to resolve enigmatic relationships among basal angiosperms, Proceedings of the National Academy of Sciences USA, vol.104, pp.19363-19368, 2007.

J. Muller, Fossil pollen records of extant angiosperms, The Botanical Review, vol.47, pp.1-142, 1981.

D. J. Nichols and H. L. Ott, Biostratigraphy and evolution of the Momipites-Caryapollenites lineage in the Early Terti-ary in the Wind River Basin, Bohemia. Palaeobotanist, vol.2, pp.52-54, 1966.

A. D. ?-pan, Rutaceae leaf fossils from the Late Oligocene (27.23 Ma) Guang River flora of northwestern Ethiopia, Review of Palaeobotany and Palynology, vol.159, pp.188-194, 2010.

J. F. Parham, P. C. Donoghue, C. J. Bell, T. D. Calway, J. J. Head et al., Sample selection bias and presence-only distribution models: Implications for background and pseudo-absence data, Ecological Applications, vol.61, pp.181-197, 2009.

K. M. Piel, How many plant species are there, where are they and at what rate are they going extinct?, Annals of the Missouri Botanical Garden, vol.49, pp.170-176, 1971.

M. ?-pole, Plant macrofossils from the Foulden Hills Diatomite (Miocene), Journal of The Royal Society of New Zealand, vol.26, pp.1-39, 1996.

?. Posada, D. ;-?-prance, G. Beentje, H. Dransfield, J. Johns et al., The tropical flora remains undercollected, Annals of the Missouri Botanical Garden, vol.25, pp.67-71, 2000.

H. ?-qian and J. Y. , An updated megaphylogeny of plants, a tool for generating plant phylogenies and an analysis of phylogenetic community structure, The Plant Journal, vol.9, pp.233-239, 2016.

?. Team, R: A language and environment for statistical computing, R Foundation for Statistical Computing, 2009.

E. Reid and M. Chandler, Catalogue of Cainozoic plants in the department of Geology, 1926.

, The Bembridge flora. British Museum (Natural History)

L. J. Revell, L. J. Harmon, and D. C. Collar, phytools: An R package for phylogenetic comparative biology (and other things), Methods in Ecology and. Evolution, vol.57, pp.217-223, 2008.

F. Ronquist, M. Teslenko, P. Mark, D. L. Ayres, A. Darling et al., MrBayes 3.2: efficient Bayesian phylogenetic inference and model choice across a large model space, Systematic Biology, vol.61, pp.539-542, 2012.

B. ?-roure, D. Baurain, and H. Philippe, Impact of missing data on phylogenies inferred from empirical phylogenomic data sets, Molecular Biology and Evolution, vol.30, pp.197-214, 2013.

M. J. Sanderson and H. B. Shaffer, Troubleshooting molecular phylogenetic analyses, Annual Review of Ecology and Systematics, vol.33, pp.49-72, 2002.

M. J. Sanderson, Inferring absolute rates of molecular evolution and divergence times in the absence of a molecular clock, Bioinformatics, vol.8, pp.301-302, 2003.

Y. Y. Shen, X. Chen, and R. W. Murphy, Assessing DNA barcoding as a tool for species identification and data quality control, PLoS ONE, vol.8, p.57125, 2013.

H. Sims, P. Herendeen, and P. Crane, New fenus of fossil Fagaceae from the Santonian, International Journal of Plant Sciences, vol.159, pp.391-404, 1998.

S. A. Smith, J. M. Beaulieu, M. Donoghue, S. A. Smith, and O. Bc, An uncorrelated relaxed-clock analysis suggests an earlier origin for flowering plants, Proceedings of the National Academy of Sciences USA, vol.107, pp.2689-2690, 2010.

S. A. Smith, J. Brown, P. Soltis, D. Soltis, V. Savolainen et al., Rate heterogeneity among lineages of tracheophytes: integration of molecular and fossil data and evidence for molecular living fossils, Proceedings of the National Academy of Sciences USA, vol.105, pp.4430-4435, 2002.

A. Stamatakis, RAxML-VI-HPC: maximum likelihood-based phylogenetic analyses with thousands of taxa and mixed models, Bioinformatics, vol.22, pp.2688-2690, 2006.

M. A. Suchard, P. Lemey, G. Baele, D. L. Ayres, and A. J. Drummond, Bayesian phylogenetic and phylodynamic data integration using BEAST 1.10, Virus Evolution, vol.4, issue.16, 2018.

G. Sun, D. L. Dilcher, H. Wang, Z. Chen, M. Takahashi et al., Hironoia fusiformis gen. et sp. nov.: A cornalean fruit from the Kamikitaba locality (Upper Cretaceous, Lower Coniacian) in northeastern Japan, Journal of Plant Research, vol.471, pp.463-473, 2002.

N. Tel-zur, S. Abbo, D. Myslabodski, and Y. Mizrahi, Modified CTAB procedure for DNA isolation from epiphytic cacti of the genera Hylocereus and Selenicereus (Cactaceae), Plant Molecular Biology Reporter, vol.17, pp.511-522, 1999.

F. Vandelook, S. B. Janssens, and R. J. Probert, Ecological niche and phylogeny explain distribution of seed mass in the central European flora, Janssens SB, Matthies D, vol.195, pp.1410-1421, 2012.

?. Vaudois-miéja, N. Wanntorp, H. E. Brooks, D. R. Nilson, T. Nylin et al., Comptes-Rendus des Seances de l'Academie des Sciences, Série 2: Mecanique-Physique, Chimie, Sciences de l'Univers, Sciences de la Terre, Phylogenies and community ecology. Annual Review of Ecology and Systematics, vol.296, pp.475-505, 1983.

C. O. Webb, D. D. Ackerly, and S. W. Kembel, Phylocom: software for the analysis of phylogenetic community structure and trait evolution, Bioinformatics, vol.24, pp.2098-2100, 2008.

W. C. Wehr and S. Manchester, Paleobotanical significance of Eocene flowers, fruits, and seeds from Republic, Washington. Washington Geology, vol.24, pp.25-27, 1996.

N. Wikstrom, V. Savolainen, and M. W. Chase, Evolution of the angiosperms: Calibrating the family tree, Proceedings of the Royal Society of London B Biological Sciences, vol.268, pp.2211-2220, 2001.

V. Wilde, Untersuchungen zur Systematik der Blattreste aus dem Mitteleozan der Grube Messel bei Darmstadt, Courier Forschungsinstitut Senckenberg, vol.115, pp.1-213, 1989.

P. Wilf, M. R. Carvalho, M. A. Gandolfo, and N. R. Cuneo, Eocene lantern fruits from Godwanan Patagonia and the early origins of Solanaceae, Science, vol.355, pp.71-75, 2017.

Y. G. Yao, C. M. Bravi, H. J. Bandelt, A. E. ;-?-zanne, D. C. Tank et al., Three keys to the radiation of angiosperms into freezing environments, Forensic Science International, vol.141, pp.89-92, 2003.

Z. Zhou, W. L. Crepet, and K. C. Nixon, The earliest fossil evidence of the Hamamelidaceae: Late Cretaceous (Turonian) inflorescences and fruits of Altingioideae, American Journal of Botany, vol.88, pp.753-766, 2001.