A. Ayouba, F. Mouacha, G. H. Learn, E. Mpoudi-ngole, J. C. Rayner et al., Ubiquitous Hepatocystis infections, but no evidence of Plasmodium falciparum-like malaria parasites in wild greater spot-nosed monkeys (Cercopithecus nictitans), International Journal for Parasitology, vol.42, pp.709-713, 2012.

J. Borner, C. Pick, J. Thiede, O. M. Kolawole, M. T. Kingsley et al., Phylogeny of haemosporidian blood parasites revealed by a multi-gene approach, Molecular Phylogenetics and Evolution, vol.94, pp.221-231, 2016.

L. Boundenga, B. Makanga, B. Ollomo, A. Gilabert, V. Rougeron et al., Haemosporidian parasites of antelopes and other vertebrates from Gabon, Central Africa. Plos One, vol.11, 2016.
URL : https://hal.archives-ouvertes.fr/hal-01960172

L. Boundenga, B. Ollomo, V. Rougeron, L. Y. Mouele, B. Mve-ondo et al., Diversity of malaria parasites in great apes in Gabon, Malaria Journal, vol.14, p.111, 2015.
URL : https://hal.archives-ouvertes.fr/hal-01957550

L. Boundenga, S. L. Perkins, B. Ollomo, V. Rougeron, E. M. Leroy et al., Haemosporidian parasites of reptiles and birds from Gabon, Central Africa, Journal of Parasitology, vol.103, pp.330-337, 2017.
URL : https://hal.archives-ouvertes.fr/hal-02005457

J. Cox-singh, M. S. Yau, B. Singh, M. T. Abdullah, I. H. Zaidi et al., Malaria-like parasites of small mammals in Malaysian Borneo (only nucleotidic sequences are available in Genbank), 2006.

A. Dereeper, V. Guignon, G. Blanc, S. Audic, S. Buffet et al., Phylogeny.fr: Robust phylogenetic analysis for the non-specialist, Nucleic Acids Research, vol.36, pp.465-469, 2008.
URL : https://hal.archives-ouvertes.fr/lirmm-00324099

L. Duval, C. Mejean, G. D. Maganga, B. K. Makanga, L. B. Mangama-koumba et al., The chiropteran haemosporidian Polychromophilus melanipherus: A worldwide species complex restricted to the family Miniopteridae, Infection, Genetics and Evolution, vol.12, pp.1558-1566, 2012.

C. Faust and A. P. Dobson, Primate malarias: Diversity, distribution and insights for zoonotic Plasmodium, One Health, vol.1, pp.66-75, 2015.

P. C. Garnham, Malaria parasites and other haemosporidia, vol.1132, 1966.

S. Guindon, F. Lethiec, P. Duroux, and O. Gascuel, PHYML online-a web server for fast maximum likelihood-based phylogenetic inference, Nucleic Acids Research, vol.33, 2005.
URL : https://hal.archives-ouvertes.fr/lirmm-00105317

H. N. Hatta, P. C. Divis, and B. Singh, Molecular characterization of haemosporidian parasites in small mammals (only nucleotidic sequences are available in Genbank), 2012.

J. P. Huelsenbeck and F. Ronquist, MRBAYES: Bayesian inference of phylogenetic trees, Bioinformatics, vol.17, pp.754-755, 2001.

G. Karadjian, A. Hassanin, B. Saintpierre, G. Gembu-tungaluna, F. Ariey et al., Highly rearranged mitochondrial genome in Nycteria parasites (Haemosporidia) from bats, Proceedings of the National Academy of Sciences USA, vol.113, pp.9834-9839, 2016.
URL : https://hal.archives-ouvertes.fr/hal-01395176

N. D. Levine, Some corrections of coccidian (Apicomplexa: Protozoa) nomenclature, Journal of Parasitology, vol.66, pp.830-834, 1980.

G. D. Maganga, M. Bourgarel, P. Vallo, T. D. Dallo, C. Ngoagouni et al., Bat distribution size or shape as determinant of viral richness in african bats, PLoS One, vol.9, 2014.

E. S. Martinsen and S. L. Perkins, The diversity of Plasmodium and other Haemosporidians: The interesection of taxonomy, phylogenetics and genomics. Malaria parasites: Comparative genomics, evolution and molecular biology, pp.1-15, 2013.

A. Megali, G. Yannic, and P. Christe, Disease in the dark: Molecular characterization of Polychromophilus murinus in temperate zone bats revealed a worldwide distribution of this malaria-like disease, Molecular Ecology, vol.20, pp.1039-1048, 2011.

K. J. Olival, E. O. Stiner, and S. L. Perkins, Detection of Hepatocystis sp. in southeast Asian flying foxes (Pteropodidae) using microscopic and molecular methods, Journal of Parasitology, vol.93, pp.1538-1540, 2007.

S. L. Perkins, Malaria's many mates: Past, present, and future of the systematics of the order Haemosporida, Journal of Parasitology, vol.100, pp.11-25, 2014.

D. Posada and K. A. Crandall, Modeltest: Testing the model of DNA substitution, Bioinformatics, vol.14, pp.817-818, 1998.

F. Prugnolle, P. Durand, C. Neel, B. Ollomo, F. J. Ayala et al., African great apes are natural hosts of multiple related malaria species, including Plasmodium falciparum, Proceedings of the National Academy of Sciences USA, vol.107, pp.1458-1463, 2010.

F. Prugnolle, B. Ollomo, P. Durand, E. Yalcindag, C. Arnathau et al., African monkeys are infected by Plasmodium falciparum nonhuman primate-specific strains, Proceedings of the National Academy of Sciences USA, vol.108, pp.11948-11953, 2011.

C. Putaporntip, S. Jongwutiwes, S. Thongaree, S. Seethamchai, P. Grynberg et al., Ecology of malaria parasites infecting Southeast Asian macaques: Evidence from cytochrome b sequences, Molecular Ecology, vol.19, pp.3466-3476, 2010.

J. Schaer, S. L. Perkins, J. Decher, F. H. Leendertz, J. Fahr et al., High diversity of West African bat malaria parasites and a tight link with rodent Plasmodium taxa, Proceedings of the National Academy of Sciences USA, vol.110, pp.17415-17419, 2013.

J. Schaer, S. L. Perkins, I. Ejotre, M. E. Vodzak, K. Matuschewski et al., Epauletted fruit bats display exceptionally high infections with a Hepatocystis species complex in South Sudan, Scientific Reports, vol.7, 2017.

J. Schaer, D. M. Reeder, M. E. Vodzak, K. J. Olival, N. Weber et al., Nycteria parasites of Afrotropical insectivorous bats, International Journal for Parasitology, vol.45, pp.375-384, 2015.

S. Seethamchai, C. Putaporntip, S. Malaivijitnond, L. Cui, and S. Jongwutiwes, Malaria and Hepatocystis species in wild macaques, southern Thailand, American Journal of Tropical Medicine and Hygiene, vol.78, pp.646-653, 2008.

M. A. Suchard, P. Lemey, G. Baele, D. L. Ayres, A. J. Drummond et al., Bayesian phylogenetic and phylodynamic data integration using BEAST 1.10. Virus Evolution, vol.4, 2018.

M. I. Thurber, R. R. Ghai, D. Hyeroba, G. Weny, A. Tumukunde et al., Co-infection and cross-species transmission of divergent Hepatocystis lineages in a wild African primate community, International Journal for Parasitology, vol.43, pp.613-619, 2013.

M. Vafa-homann, S. N. Emami, V. Yman, C. Stenström, K. Sondén et al., Detection of malaria parasites after treatment in travelers: A 12-months longitudinal study and statistical modelling analysis, EBioMedicine, vol.25, pp.66-72, 2017.

S. Fo-r-m-ati-o-n-;-boundenga, L. Ngoubangoye, B. Mombo, and I. M. , Additional supporting information may be found online in the Supporting Information section at the end of the article. How to cite this article, Ecol Evol, vol.8, pp.10578-10586, 2018.