D. C. Adams, A generalized K statistic for estimating phylogenetic signal from shape and other high-dimensional multivariate data, Syst Biol, vol.63, pp.685-697, 2014.

D. C. Adams and E. Otárola-castillo, Geomorph: an R package for the collection and analysis of geometric morphometric shape data, Methods Ecol Evol, vol.4, pp.393-399, 2013.

D. C. Adams, F. J. Rohlf, and D. Slice, Geometric morphometrics: ten years of progress following the Brevolution.^Hystrix, Ital J Zool, vol.71, pp.5-16, 2004.

K. P. Aplin, P. R. Brown, J. Jacob, C. J. Krebs, and G. R. Singleton, Field methods for rodent studies in Asia and the Indo-Pacific: review of the major pest species, ACIAR Monograph Series, issue.100, 2003.

, Australian Centre for International Agricultural Research, pp.121-223

K. P. Aplin, H. Suzuki, A. A. Chinen, R. T. Chesser, J. Ten-have et al., Multiple geographic origins of commensalism and complex dispersal history of black rats, PLoS One, vol.6, p.26357, 2011.
URL : https://hal.archives-ouvertes.fr/ird-00714383

P. L. Armitage, Unwelcome companions: ancient rats reviewed, Antiquity, vol.68, pp.231-240, 1994.

P. L. Armitage, B. West, and K. Steedman, New evidence of black rat in Roman London, Lond Archaeol, vol.4, pp.375-383, 1984.

A. D. Bastos, D. Nair, P. J. Taylor, H. Brettschneider, F. Kirsten et al., Genetic monitoring detects an overlooked cryptic species and reveals the diversity and distribution of three invasive Rattus congeners in South Africa, BMC Genet, vol.12, p.26, 2011.

S. P. Blomberg, T. J. Garland, and A. R. Ives, Testing for phylogenetic signal in comparative data: behavioral traits are more labile, Evolution, vol.57, pp.717-745, 2003.

F. L. Bookstein, Landmark methods for forms without landmarks: morphometrics of group differences in outline shape, Med Image Anal, vol.1, pp.225-243, 1997.

M. A. Butler and A. A. King, Phylogenetic comparative analysis: a modeling approach for adaptive evolution, Am Nat, vol.164, pp.683-695, 2004.

D. Capizzi, S. Bertolino, and A. Mortelliti, Rating the rat: global patterns and research priorities in impacts and management of rodent pests, Mammal Rev, vol.44, pp.148-162, 2014.

Y. Chaimanee and J. Jaeger, Evolution of Rattus (Mammalia, Rodentia) during the Plio-Pleistocene in Thailand, Hist Biol, vol.15, pp.181-191, 2001.

Y. Chaval, S. Waengsothorn, S. Morand, J. F. Cosson, and C. J. , A new taxonomic toolkit for identification of two sympatric species of Bandicota (Rodentia: Muridae) from Mainland Southeast Asia, Mammalia, vol.80, pp.425-439, 2016.
URL : https://hal.archives-ouvertes.fr/hal-01602398

D. S. Chingangbam, J. M. Laishram, and H. Suzuki, Molecular phylogenetic characterization of common murine rodents from Manipur, northeast India, Genes Genet Syst, vol.90, pp.21-30, 2015.

J. Claude, R. Morphometrics, and . Springer, Log-Shape ratios, Procrustes superimposition, elliptic Fourier analysis: three worked examples in R, hystrix. Ital Mammal, vol.24, pp.94-102, 2008.

C. J. Conroy, K. C. Rowe, K. Rowe, P. L. Kamath, K. P. Aplin et al., Cryptic genetic diversity in Rattus of the San Francisco Bay region, California. Biol Invasions, vol.15, pp.741-758, 2013.

T. Cucchi, R. Barnett, N. Martínková, S. Renaud, E. Renvoisé et al., The changing pace of insular life: 5000 years of microevolution in the Orkney vole (Microtus arvalis orcadensis), Evolution, vol.68, pp.2804-2820, 2014.
URL : https://hal.archives-ouvertes.fr/hal-02489428

T. Cucchi, Z. E. Kovács, R. Berthon, A. Orth, F. Bonhomme et al., On the trail of Neolithic mice and men towards Transcaucasia: zooarchaeological clues from Nakhchivan (Azerbaijan), Biol J Linn Soc, vol.1084, pp.917-928, 2013.
URL : https://hal.archives-ouvertes.fr/hal-02043993

G. Cuenca-bescós, L. G. Straus, G. Morales, M. R. , G. Pimienta et al., The reconstruction of past environments through small mammals: from the Mousterian to the Bronze Age in El Mirón Cave (Cantabria Spain), J Archaeol Sci, vol.36, pp.947-955, 2009.

T. Dayan and D. Simberloff, Character displacement, sexual dimorphism, and morphological variation among British and Irish mustelids, Ecology, vol.75, pp.1063-1073, 1994.

T. Dayan and D. Simberloff, Size patterns among competitors: ecological character displacement and character release in mammals with special reference to island populations, Mammal Rev, vol.28, pp.99-124, 1998.

T. Dayan, D. Simberloff, E. Tchernov, and Y. Yom-tov, Inter-and intraspecific character displacement in mustelids, Ecology, vol.70, pp.1526-1539, 1989.

T. Dayan, D. Simberloff, E. Tchernov, and Y. Yom-tov, Canine carnassials: character displacement in the wolves, jackals and foxes of Israel, Biol J Linn Soc, vol.45, pp.315-331, 1992.

T. Dayan, E. Tchernov, Y. Yom-tov, and D. Simberloff, Ecological character displacement in Saharo-Arabian Vulpes: outfoxing Bergmann's rule, Oikos, vol.55, pp.263-272, 1989.

A. Evin, T. Cucchi, A. Cardini, U. Strand-vidarsdottir, G. Larson et al., The long and winding road: identifying pig domestication through molar size and shape, J Archaeol Sci, vol.40, pp.735-743, 2013.

P. Fabre, L. Hautier, D. Dimitrov, and E. Douzery, A glimpse on the pattern of rodent diversification: a phylogenetic approach, BMC Evol Biol, vol.12, pp.1-19, 2012.

J. Felsenstein, Maximum-likelihood estimation of evolutionary trees from continuous characters, Am J Hum Genet, vol.25, pp.471-492, 1973.

L. J. Harmon and R. E. Glor, Poor statistical performance of the Mantel test in phylogenetic comparative analyses, Evolution, vol.64, pp.2173-2178, 2010.

L. J. Harmon, J. B. Losos, T. J. Davies, R. G. Gillespie, J. L. Gittleman et al., Early bursts of body size and shape evolution are rare in comparative data, Evolution, vol.64, pp.2385-2396, 2010.

L. J. Harmon, J. T. Weir, C. D. Brock, R. E. Glor, and W. Challenger, GEIGER : investigating evolutionary radiations, Bioinformatics, vol.24, pp.129-131, 2008.

L. Hautier, P. Fabre, and J. Michaux, Mandible shape and dwarfism in squirrels (Mammalia Rodentia): interaction of allometry and adaptation, Naturwissenschaften, vol.96, pp.725-730, 2009.

C. P. Klingenberg and N. A. Gidaszewski, Testing and quantifying phylogenetic signals and homoplasy in morphometric data, Syst Biol, vol.59, pp.245-261, 2010.

M. Kosoy, L. Khlyap, J. Cosson, and S. Morand, Aboriginal and invasive rats of genus Rattus as hosts of infectious agents, Vector-Borne Zoonotic Dis, vol.15, pp.3-12, 2015.

J. B. Lack, D. U. Greene, C. J. Conroy, M. J. Hamilton, J. K. Braun et al., Invasion facilitates hybridization with introgression in the Rattus rattus species complex, Mol Ecol, vol.21, pp.3545-3561, 2012.

E. Lecompte, K. P. Aplin, C. Denys, F. Catzeflis, M. Chades et al., Phylogeny and biogeography of African Murinae based on mitochondrial and nuclear gene sequences with a new tribal classification of the subfamily, BMC Evol Biol, vol.8, p.199, 2008.
URL : https://hal.archives-ouvertes.fr/hal-00336780

M. A. Mares, South American mammal zoogeography: evidence from convergent evolution in desert rodents, Proc Natl Acad Sci, vol.72, pp.1702-1706, 1975.

S. Meiri, N. Cooper, and A. Purvis, The island rule: made to be broken?, Proc Biol Sci, vol.275, pp.141-148, 2008.

S. Meiri, T. Dayan, and T. Aviv, On the validity of Bergmann's rule, J Biogeogr, vol.30, pp.331-351, 2003.

S. Meiri, P. Raia, and A. B. Phillimore, Slaying dragons: limited evidence for unusual body size evolution on islands, J Biogeogr, vol.38, pp.89-100, 2011.

C. Meloro and P. Raia, Cats and dogs down the tree: the tempo and mode of evolution in the lower carnassial of fossil and living Carnivora, Evol Biol, vol.37, pp.177-186, 2010.

V. Millien and J. Damuth, Climate change and size evolution in an island rodent species: new perspectives on the island rule, Evolution, vol.58, pp.1353-1360, 2004.

V. Millien, S. K. Lyons, L. Olson, F. A. Smith, A. B. Wilson et al., Ecotypic variation in the context of global climate change: revisiting the rules, Ecol Lett, vol.9, pp.853-869, 2006.

P. Mitteroecker, P. Gunz, M. Bernhard, K. Schaefer, and F. L. Bookstein, Comparison of cranial ontogenetic trajectories among great apes and humans, J Hum Evol, vol.46, pp.679-697, 2004.

S. Morand, F. Bordes, H. Chen, C. J. Cosson, J. Galan et al., Global parasite and Rattus rodent invasions: the consequences for rodentborne diseases, Integr Zool, vol.10, pp.409-423, 2013.
URL : https://hal.archives-ouvertes.fr/hal-01920147

S. Morand, F. Bordes, K. Blasdell, S. Pilosof, J. Cornu et al., Assessing the distribution of disease-bearing rodents in human-modified tropical landscapes, J Applied Ecol, vol.52, pp.784-794, 2015.
URL : https://hal.archives-ouvertes.fr/hal-01239624

G. G. Musser, Identities of taxa associated with Rattus rattus (Rodentia, Muridae) of Sumba Island, Indonesia. J Mammal, vol.53, pp.861-865, 1972.

G. G. Musser, The giant rats of Flores and its relatives east of Borneo and Bali, Bull Am Mus Nat Hist, vol.169, pp.67-176, 1981.

G. G. Musser, Sundaic Rattus: definitions of Rattus baluensis and Rattus korinchi, Am Mus Novitates, vol.2862, pp.1-24, 1986.

G. G. Musser and C. Newcomb, Malaysian murids and the giant rat of Sumatra, Bull Am Mus Nat Hist, vol.174, pp.327-598, 1983.

N. Myers, R. A. Mittermeier, C. G. Mittermeier, G. A. Da-fonseca, and J. Kent, Biodiversity hotspots for conservation priorities, Nature, vol.403, pp.853-858, 2000.

J. Oksanen, F. G. Blanchet, M. Friendly, R. Kindt, P. Legendre et al., , 2017.

M. Pagel, Inferring the historical patterns of biological evolution, Nature, vol.401, pp.877-884, 2000.

M. Pagès, Y. Chaval, V. Herbreteau, S. Waengsothorn, J. Cosson et al., Revisiting the taxonomy of the Rattini tribe: a phylogeny-based delimitation of species boundaries, BMC Evol Biol, vol.10, p.184, 2010.

M. Pagès, G. Corbet, A. Orth, V. Volobouev, J. Michaux et al., Morphological chromosomal and genic differences between sympatric Rattus rattus and Rattus satarae in South India, J Mammal, vol.92, pp.659-670, 2011.

E. Paradis, J. Claude, and K. Strimmer, APE: analyses of phylogenetics and evolution in R language, Bioinformatics, vol.20, pp.289-290, 2004.
URL : https://hal.archives-ouvertes.fr/ird-01887318

M. J. Pearch, S. Bumrungsri, J. Schwenninger, D. J. Ward, and D. L. Harrison, A review of the Cainozoic small mammal fauna of Thailand with new records (Chiroptera ; Scandentia, 2013.

, Eulipotyphla) from the late Pleistocene. Cainozoic Res, vol.10, pp.59-98

O. Pergams, D. Byrn, K. Lee, and R. Jackson, Rapid morphological change in black rats (Rattus rattus) after an island introduction, PeerJ, vol.3, p.812, 2015.

S. Pimm, J. A. Drake, H. A. Mooney, F. Di-castri, R. H. Groves et al., Theories of predicting success and impact of introduced species, Biological Invasions: A Global Perspective, pp.351-367, 1989.

P. Piras, C. J. Ventura, J. Cubo, J. Pierre, U. Paris et al., Ecological and functional correlates of molar shape variation in European populations of Arvicola (Arvicolinae Rodentia), Zool Anzeiger, vol.251, pp.335-343, 2012.
URL : https://hal.archives-ouvertes.fr/hal-01435016

M. E. Prendergast, M. Buckley, A. Crowther, L. Frantz, H. M. Eager et al., Reconstructing Asian faunal introductions to eastern Africa from multi-proxy biomolecular and archaeological datasets, PLoS One, vol.12, issue.8, p.182565, 2017.

. R-core-team, R: A language and environment for statistical computing. R Foundation for Statistical Computing, 2016.

S. Renaud, S. Pantalacci, and J. C. Auffray, Differential evolvability along lines of least resistance of upper and lower molars in island house mice, PLoS One, vol.6, issue.5, p.18951, 2011.
URL : https://hal.archives-ouvertes.fr/hal-00698157

L. J. Revell, phytools: an R package for phylogenetic comparative biology (and other things), Methods Ecol Evol, vol.3, pp.217-223, 2012.

J. H. Robins, M. Hingston, E. Matisoo-smith, and H. A. Ross, Identifying Rattus species using mitochondrial DNA, Mol Ecol Notes, vol.7, pp.717-729, 2007.

J. H. Robins, V. Tintinger, K. P. Aplin, M. Hingston, E. Matisoo-smith et al., Phylogenetic species identification in Rattus highlights rapid radiation and morphological similarity of New Guinean species, PLoS One, vol.9, issue.5, p.98002, 2014.

F. J. Rohlf, Recent and rapid speciation with limited morphological disparity in the genus Rattus, TpsRelw 1.41-Thin plate Spline Relative Warp, 1.41, vol.60, pp.188-203, 2005.

K. C. Rowe, M. L. Reno, D. M. Richmond, R. M. Adkins, and S. J. Steppan, Pliocene colonization and adaptive radiations in Australia and New Guinea (Sahul): multilocus systematics of the old endemic rodents (Muroidea: Murinae), vol.47, pp.84-101, 2008.

M. A. Serrat, R. M. Williams, and C. E. Farnum, Exercise mitigates the stunting effect of cold temperature on limb elongation in mice by increasing solute delivery to the growth plate, J Appl Physiol, vol.109, pp.1869-1879, 2010.

A. B. Shiels, C. A. Flores, A. Khamsing, P. D. Krushelnycky, S. M. Mosher et al., Dietary niche differentiation among three species of invasive rodents (Rattus rattus R exulans Mus musculus), Biol Invasions, vol.15, pp.1037-1048, 2012.

E. Stoetzel, L. Marion, R. Nespoulet, M. Abdeljalil, E. Hajraoui et al., Taphonomy and palaeoecology of the late Pleistocene to middle Holocene small mammal succession of El Harhoura 2 cave (Rabat-Témara Morocco), J Hum Evol, vol.60, pp.1-33, 2011.

J. M. Taylor, J. H. Calaby, and H. M. Van-deusen, A revision of the genus Rattus (Rodentia Muridae) in the New Guinean region, Bull Am Mus Nat Hist, vol.173, pp.179-336, 1982.

J. W. Tukey, Comparing individual means in the analysis of variance, Biometrics, vol.5, pp.99-114, 1949.

L. Weissbrod, F. B. Marshall, F. R. Valla, H. Khalaily, G. Bar-oz et al., Origins of house mice in ecological niches created by settled hunter-gatherers in the Levant 15000 y ago, Proc Natl Acad Sci, vol.114, pp.1-6, 2017.
URL : https://hal.archives-ouvertes.fr/mnhn-02363694

A. E. Wood, Grades and clades among rodents, Evolution, vol.19, pp.115-130, 1965.

Y. Yom-tov, Y. , and S. , Climatic change and body size in two species of Japanese rodents, Biol J Linn Soc, vol.82, pp.263-267, 2004.