A fundamental and long-standing question of southern African palaeoclimatology is the way tropical and temperate climate system dynamics have influenced rainfall regimes across the subcontinent since the Last glacial maximum. In this paper, we analyse a selection of recently published palaeoclimate reconstructions along a southwest-northeast transect across South Africa. These records span the last 22,000 years, and encompass the transition between the region's winter and summer rainfall zones. In synthesis, these records confirm broad elements of the dominant paradigm, which proposes an inverse coeval relationship between temperate and tropical systems, with increased precipitation in the winter (summer) rainfall zone during glacial (interglacial) periods. Revealed, however, is a substantially more complex dynamic, with millennial-scale climate change events being strongly e even predominantly e influenced by the interaction and combination of temperate and tropical systems. This synoptic forcing can create same sign anomalies across the South African rainfall zones, contrary to expectations based on the classic model of phase opposition. These findings suggest a new paradigm for the interpretation of southern African palaeoenvironmental records that moves beyond simple binary or additive influences of these systems.